Biology 325: Advanced Genetics

Lecture I

Outline:           I.          Introduction

II.             Early Studies of DNA

III.           Molecular Structure of DNA

IV.          Genetic Implications of DNA Structure

V.            Special Structures in RNA and DNA

VI.          DNA Methylation

VII.        RNA as Genetic Material

 

 

I.               Introduction: The Story of the Ice Man

 

A.   Discovery

 

 

 

 

B.    Questions

 

 

 

 

C.    DNA Evidence

 

 

 

 

 

 

II.             Early Studies of DNA

 

A.   Fredrich Miescher (1869)

 

 

 

B.    Albrecht Kessel

 

 

 

C.    Pheobus Aaron Levene

 

 

 

D.   Erwin Chargaff (1948)

 

III.           Molecular Structure of DNA

 

A.   Rosalind Franklin and Maurice Wilkins

 

 

 

 

B.    James D. Watson and Francis Crick (1953)

 

 

 

 

 

 

 

C.    The Double Helix

 

1.     Components

 

 

 

 

 

2.     Nucleosides

 

 

 

 

 

 

 

3.     Nucleotide

 

 

 

 

 

 

4.     Polynucleotide Strands

 

 

 

 

 

 

 

5.     Double-stranded DNA

 

a.     B-DNA

 

 

 

 

b.     A-DNA

 

 

 

c.     Z-DNA

 

 

 

Characteristic

B-DNA

A-DNA

Z-DNA

Conditions required to produce structure

92% H20

75% H20

Alternating purine-pyrmidines

Helix Direction

Right-handed

Right-handed

Left-handed

Average base pairs per turn

10

11

12

Rotation per base pair

    36¡

32.7¡

-30¡

Distance between bases

0.34nm

0.26nm

0.37nm

Diameter of helix

1.9nm

2.3nm

1.8nm

Overall shape

Long and narrow

Short and wide

Elongated and narrow

 

IV.          Genetic Implications of DNA Structure

 

A.   Central Dogma of Genetics

 

 

 

 

 

 

 

 

 

 

 

V.            Special Structures in RNA and DNA

 

A.   Base pairing within single stranded polynucleotide strands

 

 

 

 

B.    Hairpins, Stems, and Cruciforms

 

 

 

 

 

 

 

 

VI.          DNA Methylation

 

A.   DNA modification

 

 

 

 

B.    Function of Methylation

 

 

 

 

 

 

VII.        RNA as Genetic Material

 

A.   Fraenkal-Conrad and Singer (1956)

 

 

 

 

 

 

 

 

B.    Central Dogma in Retroviruses